I gave the model a look and found some group comment discrepancies which I fixed, they might have been the problem, or not.
I also made some changes in the names of some groups for my own convenience (I needed to know what each group was without the need to look at the 3D viewport).
The thing, however, that hit me right away was the lod1 designation in the name of the model.
If you are only going to use one lod, then it must be lod0.
Other than that, you need to make sure that your model is using the correct animations.
That is a matter of scripting.
Here is my version with small fixes, let me know if it works.
Marchwardens_Fix-1.7z